{"id":27830,"date":"2021-07-28T23:56:24","date_gmt":"2021-07-28T15:56:24","guid":{"rendered":"https:\/\/swarma.org\/?p=27830"},"modified":"2021-07-28T23:56:24","modified_gmt":"2021-07-28T15:56:24","slug":"science%e5%b0%81%e9%9d%a2%e6%96%87%e7%ab%a0%e8%a7%a3%e8%af%bb%ef%bc%9a%e8%8a%b1%e6%a4%b0%e8%8f%9c%e5%88%86%e5%bd%a2%e7%bb%93%e6%9e%84%e7%9a%84%e8%b5%b7%e6%ba%90","status":"publish","type":"post","link":"https:\/\/swarma.org\/?p=27830","title":{"rendered":"Science\u5c01\u9762\u6587\u7ae0\u89e3\u8bfb\uff1a\u82b1\u6930\u83dc\u5206\u5f62\u7ed3\u6784\u7684\u8d77\u6e90"},"content":{"rendered":"
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Cauliflower fractal forms arise from perturbations of floral gene networks View ORCID Profile<\/span><\/p>\n

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https:\/\/science.sciencemag.org\/content\/373\/6551\/192?rss=1<\/span><\/p>\n<\/blockquote>\n


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4.\u7f57\u9a6c\u82b1\u6930\u83dc\u82b1\u7403\u7684\u52a8\u6001\u53d1\u5c55\u6a21\u578b<\/span><\/strong><\/span><\/section>\n

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    \u8fdb\u4e00\u6b65\u7814\u7a76\u53d1\u73b0\uff0c\u82b1\u6570\u3001\u53f6\u5e8f\u548c\u679c\u5b9e\u7684\u87ba\u65cb\u7ebf\u5728\u751f\u957f\u8fc7\u7a0b\u4e2d\u6240\u63a5\u8fd1\u7684\u89d2\u5ea6\u5c31\u662f 1\/(1+\u03d5\uff09\uff0c\u5bf9\u5e94\u5706\u5468\u89d2\u5927\u7ea6137.507764\u2026\u2026\u00b0\u7684\u503c\uff0c\u5b83\u4e5f\u88ab\u79f0\u4e3a\u9ec4\u91d1\u89d2\u5ea6<\/strong>\u3002\u539f\u57fa\u5728\u690d\u7269\u830e\u5c16\u7684\u751f\u957f\u89c4\u5f8b\u5982\u56fe2\u6240\u793a\uff0c\u6700\u7ec8\u5404\u4e2a\u539f\u57fa\u5448\u87ba\u65cb\u72b6\u4ee5\u9ec4\u91d1\u89d2\u5ea6\u7684\u500d\u6570\u4f9d\u6b21\u6392\u5f00\uff0c\u5f62\u6210\u4e86\u830e\u53f6\u76f8\u95f4\u7684\u5947\u7279\u7684\u51e0\u4f55\u6837\u8c8c\u3002<\/span><\/section>\n
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    \u4e0d\u8981\u5c0f\u770b\u9ec4\u91d1\u89d2\u5ea6\u7684\u7cbe\u786e\u503c\uff0c\u53ea\u6709\u6cbf137.5\u00b0\u751f\u957f\uff0c\u624d\u4f1a\u5f62\u6210\u56fe3\u4e2d\u95f4\u7684\u8d39\u9a6c\u87ba\u7ebf<\/strong>\u3002\u6b64\u5916\u54ea\u6015\u6709\u5341\u5206\u4e4b\u4e00\u7684\u504f\u5dee\uff0c\u90fd\u4f1a\u5f62\u6210\u622a\u7136\u4e0d\u540c\u7684\u7ed3\u679c[2]\u3002\u5728\u6cbf\u8d39\u9a6c\u87ba\u7ebf\u65f6\uff0c\u5411\u65e5\u8475\u79cd\u5b50\u4f1a\u7d27\u5bc6\u805a\u96c6\u5728\u4e00\u8d77\uff0c\u4e2d\u95f4\u6ca1\u6709\u7f1d\u9699\u6216\u91cd\u53e0\u3002\u6b64\u65f6\u79cd\u5b50\u5145\u5206\u5229\u7528\u4e86\u7a7a\u95f4\uff0c\u5806\u79ef\u6548\u7387\u6700\u9ad8<\/strong>\u3002<\/span><\/section>\n
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    \u4e0d\u4ec5\u89d2\u5ea6\uff0c\u5177\u6709\u87ba\u65cb\u53f6\u5e8f\u7684\u690d\u7269\u5668\u5b98\uff0c\u8fd8\u4f1a\u901a\u5e38\u5f62\u6210\u4e24\u4e2a\u87ba\u65cb\u7c07\u65cf<\/span>\uff08\u56fe1A-C\uff09<\/span>\uff0c\u4ee5\u6b63\u53cd\u65b9\u5411\u8f6c\u52a8\uff0c\u5e76\u51fa\u73b0\u5728\u6590\u6ce2\u90a3\u5951\u6570\u5217\u7684\u4e24\u4e2a\u8fde\u7eed\u6570\u5b57\u4e2d[1]\u3002\u4f8b\u5982\u96cf\u83ca<\/span>\uff08\u56fe4A\uff09<\/span>\u6216\u5411\u65e5\u8475\u82b1\u76d8\u4e0a\u7684\u79cd\u5b50\uff0c\u9006\u65f6\u6216\u987a\u65f6\u9488\u87ba\u65cb\u7ebf\u7684\u6570\u91cf\uff0c\u5f80\u5f80\u90fd\u662f8\u300113\u300121 \u6216 34\u6761\u3002<\/span><\/section>\n

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    \u56fe4\uff1a(A) \u96cf\u83ca\u5934\u72b6\u82b1\u5e8f\uff0c13\u4e2a\u84dd\u8272\u987a\u65f6\u9488\u548c21\u4e2a\u9006\u65f6\u9488\u7ea2\u8272\u87ba\u65cb (B) \u5927\u4e3d\u82b1\u590d\u5408\u82b1 (C) \u751f\u59dc (D-F) \u82b1\u6930\u83dc\uff0c\u516b\u4e2a\u9006\u65f6\u9488\uff08\u68d5\u8272\uff09\u548c\u4e94\u4e2a\u987a\u65f6\u9488\uff08\u9752\u8272\uff09\u7684\u87ba\u65cb\u3002\u7ebf\u77e9\u5f62\u5c55\u793a\u4e86\u5d4c\u5957\u5728\u591a\u4e2a\u5c3a\u5ea6\u4e0a\u7684\u87ba\u65cb\u5e8f\u5217 (G) \u7f57\u9a6c\u82b1\u6930\u83dc\u82b1\u7403 (H) \u91ce\u751f\u578b (WT) \u62df\u5357\u82a5\u82b1\u5e8f\uff0c\u53ca (I) ap1 cal \u53d8\u79cd\u82b1\u7403\u3002<\/span><\/p>\n

     <\/span><\/section>\n
    \u800c\u5728\u82b1\u6930\u83dc<\/strong>\u4e2d\uff0c\u4e0d\u4ec5\u4e00\u4e2a\u5c3a\u5ea6\uff0c\u5728\u591a\u4e2a\u5c3a\u5ea6\u4e0a\u90fd\u53ef\u4ee5\u770b\u5230\u51dd\u4e73\u72b6\u7684\u87ba\u65cb\u7c07\u65cf<\/span>\uff08\u56fe4D-F\uff09<\/span>\uff0c\u6bcf\u4e2a\u82b1\u83dc\u7684\u5c0f\u82b1\u7403\u5c31\u50cf\u662f\u7531\u4e00\u4e2a\u4e2a\u5fae\u578b\u7248\u672c\u7684\u82b1\u83dc\u7ec4\u6210\u7684\u3002\u8fd9\u79cd\u7ec4\u7ec7\u81ea\u76f8\u4f3c\u6027<\/strong>\u5728\u7f57\u9a6c\u82b1\u6930\u83dc<\/strong><\/span>\uff08Romanesco\uff09<\/span>\u4e2d\u8fbe\u5230\u4e86\u9876\u5cf0\uff0c\u5176\u87ba\u65cb\u5448\u73b0\u7c7b\u4f3c\u4e8e\u6d6e\u96d5\uff0c\u5728\u6240\u6709\u5c3a\u5ea6\u4e0a\u90fd\u662f\u5706\u9525\u5f62\uff0c\u6574\u4e2a\u51e0\u4f55\u5916\u89c2\u8d4b\u4e88\u4e86\u82b1\u7403\u660e\u663e\u7684\u5206\u5f62\u7279\u5f81<\/span>\uff08\u56fe4G\uff09<\/span>\u3002<\/span><\/section>\n
     <\/span><\/section>\n
    \u5982\u4f55\u89e3\u91ca\u690d\u7269\u61c2\u6570\u5b66\u7684\u73b0\u8c61\uff1f<\/span><\/section>\n
     <\/span><\/section>\n
    \u7ef4\u591a\u5229\u4e9a\u65f6\u4ee3\u7684\u79d1\u5b66\u5bb6\u4eec\u4fe1\u8a93\u65e6\u65e6\u5730\u8ba4\u4e3a\uff0c\u7531\u4e8e\u9ec4\u91d1\u5206\u5272\u662f\u65e0\u7406\u6570\uff0c\u82b1\u53f6\u7684\u504f\u5dee\u89d2\u5ea6\u6216\u5668\u5b98\u87ba\u65cb\uff0c\u4e0d\u4ec5\u8ba9\u7a7a\u95f4\u5229\u7528\u6548\u7387\u6700\u9ad8\uff0c\u800c\u4e14\u80fd\u907f\u514d\u5668\u5b98\u4e4b\u95f4\u4e92\u76f8\u906e\u76d6\uff0c\u7531\u6b64\u624d\u80fd\u66f4\u6709\u6548\u5730\u63a5\u6536\u96e8\u6c34\u548c\u9633\u5149\u751f\u957f\u3002<\/strong><\/span><\/section>\n
     <\/span><\/section>\n
    \u4f46\u8fd9\u79cd\u7b80\u5355\u89e3\u91ca\uff0c\u5c31\u597d\u50cf\u672a\u535c\u5148\u77e5\u5730\u9884\u8bbe\u4e86\u690d\u7269\u4eec\u751f\u957f\u7684\u76ee\u7684\uff0c\u8ba4\u4e3a\u690d\u7269\u559c\u6b22\u6590\u6ce2\u90a3\u5951\u6570\u5217\u6216\u5206\u5f62\u662f\u539f\u56e0\u4e00\u6837\uff0c\u5374\u5e76\u6ca1\u6709\u7ed9\u51fa\u300c\u690d\u7269\u4e3a\u4ec0\u4e48\u61c2\u6570\u5b66\u300d\u7684\u5f62\u6210\u673a\u5236\u3002\u7b54\u6848\u5f53\u7136\u4e0d\u4f1a\u8fd9\u4e48\u7b80\u5355\u3002<\/span><\/section>\n
     <\/span><\/section>\n
    \u6700\u8fd1\uff0c\u6765\u81ea\u6cd5\u56fd\u690d\u7269\u5b66\u5bb6\u548c\u6570\u5b66\u5bb6\u5728 Nature<\/em>\u4e0a\u53d1\u5e03\u7684\u4e00\u9879\u7814\u7a76\u300aCauliflower fractal forms arise from perturbations of floral gene networks View ORCID Profile\u300b\uff0c\u5bf9\u901a\u8fc7\u7814\u7a76\u62df\u5357\u82a5\u548c\u82b1\u6930\u83dc\u82b1\u7403\u5f62\u6210\u673a\u5236\uff0c\u5efa\u7acb\u4e86\u4e00\u4e2a\u57fa\u4e8e\u57fa\u56e0\u8c03\u63a7\u7f51\u7edc<\/strong><\/span> (GRN) <\/span>\u548c\u5f62\u6001\u52a8\u529b\u5b66\u53c2\u6570\u7684\u6a21\u578b\uff0c\u4ece\u5206\u5b50\u5c42\u6b21\u89e3\u91ca\u4e86\u8c03\u63a7\u7f57\u9a6c\u82b1\u6930\u83dc\u82b1\u7403\u7684\u5f62\u6210\u5177\u6709\u660e\u663e\u5206\u5f62\u6570\u5b66\u7279\u5f81\u7684\u539f\u56e0\uff0c\u5bf9\u8fd9\u4e00\u6a2a\u8de8\u6570\u767e\u5e74\u7684\u8c1c\u9898\u7ed9\u51fa\u4e86\u4e00\u4e2a\u590d\u6742\u4f46\u786e\u5207\u7684\u56de\u7b54\u3002<\/span><\/section>\n
     <\/span><\/section>\n

    <\/section>\n

    \n
    \n
    \n
    \n
    \n

    <\/section>\n

    <\/section>\n
    \n
    <\/svg><\/section>\n<\/section>\n<\/section>\n
    \n

    2. \u6784\u5efa\u690d\u7269\u53d1\u80b2\u7684SALT\u57fa\u56e0\u8c03\u63a7\u7f51\u7edc<\/strong><\/strong><\/p>\n<\/section>\n

    \n

    <\/section>\n

    <\/section>\n<\/section>\n<\/section>\n<\/section>\n<\/section>\n<\/h3>\n

    <\/section>\n
    \u82b1\u6930\u83dc\u4ece\u5377\u5fc3\u83dc\u9a6f\u5316\u800c\u6765\uff0c\u82b1\u5e8f\u5448\u87ba\u65cb\u72b6\uff0c\u6bcf\u4e2a\u65b0\u51fa\u73b0\u7684\u82b1\u539f\u57fa\u6c38\u8fdc\u4e0d\u4f1a\u6210\u719f\u5230\u82b1\u671f\uff0c\u56e0\u6b64\u4f1a\u4e0d\u65ad\u4ea7\u751f\u66f4\u591a\u51dd\u4e73\u72b6\u7684\u82b1\u7403<\/strong>[4,5]\u3002\u8fd9\u79cd\u7ed3\u6784\u7531\u5341\u5b57\u82b1\u79d1\u62df\u5357\u82a5<\/strong><\/span>\uff08Arabidopsis thaliana\uff09<\/span>\u7684\u57fa\u56e0 AP1\u548cCAL \u53cc\u7a81\u53d8\u5f15\u8d77<\/span>\uff08\u56fe4H-I\uff09<\/span>\uff0c\u662f\u7f16\u7801MADS\u76d2\u86cb<\/strong>\u767d\u8f6c\u5f55\u56e0\u5b50\u3001\u4fc3\u8fdb\u82b1\u5349\u53d1\u80b2[8,9]\u7684\u4e24\u4e2a\u65c1\u7cfb\u540c\u6e90\u57fa\u56e0\u3002<\/span>
    <\/section>\n
     <\/span><\/section>\n
    \u63a7\u5236\u62df\u5357\u82a5\u82bd\u548c\u82b1\u53d1\u80b2\u7684<\/span>\u57fa\u56e0\u8c03\u63a7\u7f51\u7edc (GRN) <\/span><\/strong><\/span>\u5df2\u88ab\u5927\u91cf\u7814\u7a76\uff0c\u4f46\u80fd\u5426\u591f\u89e3\u91ca AP1 CAL \u7a81\u53d8\u5c1a\u4e0d\u6e05\u695a[10-16]\u3002\u5728\u62df\u5357\u82a5\u4e2d\uff0c\u82b1\u7684\u53d1\u80b2\u7531 LFY \u542f\u52a8<\/span>\uff08\u56fe5\uff09<\/span>\uff0cLFY \u7531 SOC1 \u548c AGL24 \u4e24\u79cdMADS\u76d2\u86cb\u767d\uff0c\u901a\u8fc7\u73af\u5883\u548c\u5185\u6e90\u6027\u7ebf\u7d22\u5728\u6574\u4e2a\u82b1\u5e8f\u5206\u751f\u7ec4\u7ec7\u4e2d\u8c03\u63a7\uff0c\u5e76\u901a\u8fc7\u6807\u8bb0\u82b1\u5206\u751f\u7ec4\u7ec7\u8d77\u59cb\u4f4d\u70b9\u7684\u690d\u7269\u751f\u957f\u7d20\u6700\u5927\u503c\u8fdb\u884c\u8bf1\u5bfc\uff0c\u4f46\u5b83\u540c\u65f6\u53c8\u88ab TFL1 \u82b1\u5e8f\u8eab\u4efd\u86cb\u767d\u6291\u5236\u3002<\/span><\/section>\n
     <\/span><\/section>\n
    \u6b64\u540e\u5728\u82b1\u539f\u57fa\u4e2d\uff0cLFY \u8bf1\u5bfc AP1 \u548c CAL\uff0c\u540c\u65f6\u5b83\u4eec\u4e5f\u5bf9 LFY \u4ea7\u751f\u6b63\u53cd\u9988<\/strong>\uff0c\u5e76\u6291\u5236SOC1\/AGL24 \u548c TFL1\uff0c\u4ece\u800c\u786e\u4fdd\u6bcf\u4e2a\u65b0\u5206\u751f\u7ec4\u7ec7\u7684\u547d\u8fd0\u662f\u6210\u4e3a\u4e00\u6735\u82b1\u3002<\/span><\/section>\n
     <\/span><\/section>\n
    <\/section>\n
    \u56fe5\uff1a\u62df\u5357\u82a5\u57fa\u56e0\u8c03\u63a7\u7f51\u7edc<\/span><\/section>\n
     <\/span><\/section>\n
    \u4f46\u5728 AP1 CAL\u7a81\u53d8\u4f53\u5982\u82b1\u6930\u83dc\u4e2d\uff0cAP1\/LFY \u6b63\u53cd\u9988\u4e0d\u5b58\u5728<\/strong>\uff0cTFL1 \u4e0d\u53d7\u4ea7\u751f\u82b1\u7684\u5206\u751f\u7ec4\u7ec7\u4e2d AP1\/CAL \u7684\u6291\u5236\u3002\u56e0\u6b64\uff0c\u5e7c\u82b1\u539f\u57fa\u4e0d\u80fd\u7ef4\u6301 LFY \u8868\u8fbe\u5e76\u5f00\u59cb\u8868\u8fbe TFL1\u3002\u7ed3\u679c\uff0c\u5b83\u4eec\u5931\u53bb\u4e86\u82b1\u6735\u8eab\u4efd\u800c\u6210\u4e3a\u522b\u6837\u82b1\u5e8f\u7684\u5206\u751f\u7ec4\u7ec7<\/strong>[8]\u3002<\/strong><\/span><\/section>\n
     <\/span><\/section>\n
    TFL1 \u5728\u65b0\u751f\u82b1\u539f\u57fa\u4e2d\u7684\u6291\u5236\u5728\u4ee5\u5f80\u7814\u7a76\u5df2\u7406\u89e3\uff0c\u4f46\u76f4\u63a5\u5bfc\u81f4\u5b83\u5728 AP1 CAL \u82b1\u5e8f\u5206\u751f\u7ec4\u7ec7\u4e2d\u4e0a\u8c03\u7684\u56e0\u7d20\u5c1a\u4e0d\u6e05\u695a\u3002\u4f5c\u8005\u901a\u8fc7\u641c\u7d22\u4e86\u9664 LFY \u4e4b\u5916\u7684 TFL1 \u7684\u76f4\u63a5\u6b63\u5411\u8c03\u8282\u56e0\u5b50\uff0c\u68c0\u67e5\u4e86 SOC1 \u548c AGL24\uff0c\u901a\u8fc7\u529f\u80fd\u6027\u4e27\u5931\u548c\u83b7\u5f97\u5b9e\u9a8c\u8868\u660e\u4e8c\u8005\u5747\u8bf1\u5bfc TFL1<\/span>\uff08\u56fe6A-I\uff09<\/span>\uff0c\u67d3\u8272\u8d28\u514d\u75ab\u6c89\u6dc0<\/span> (ChIP) <\/span>\u4e5f\u8868\u660e\u8fd9\u4e24\u4e2a\u8f6c\u5f55\u56e0\u5b50\u4f1a\u4e0e TFL1 \u533a\u57df\u7ed3\u5408\uff0c\u8c03\u8282\u5206\u751f\u7ec4\u7ec7\u4e2d\u7684\u8868\u8fbe[22]<\/span>\uff08\u56fe6J-L\uff09<\/span>\u3002\u901a\u8fc7\u5728\u8fc7\u6e21\u76f8\u8bd5\u9a8c<\/span>\uff08transient assay\uff09<\/span>\u4e2d\u6784\u5efa SOC1 \u548c AGL24\uff0c\u8868\u660e\u8fd9\u4e9b\u533a\u57df\u8db3\u4ee5\u6fc0\u6d3bTFL1 \u5e76\u4ea7\u751f\u57fa\u56e0\u62a5\u544a<\/span>\uff08\u56fe6M&N\uff09<\/span>\uff0c\u4ece\u800c\u8bc1\u5b9e\u5b83\u4eec\u662f TFL1 \u7684\u76f4\u63a5\u8c03\u8282\u5668\u3002<\/span><\/section>\n
     <\/span><\/section>\n
    <\/section>\n
    \u56fe6\uff1aTLF1\u3001SOC1 \u548c AGL24\u4f5c\u4e3a\u6b63\u5411\u8c03\u63a7\u56e0\u5b50\u7684\u5b9e\u9a8c\u7814\u7a76<\/span><\/section>\n
     <\/span><\/section>\n
    \u6b64\u5916\u56e0\u4e3a\u4e8c\u8005\u7684\u540c\u7cfb\u7269 XAL2 \u4e5f\u7ed3\u5408\u5e76\u8bf1\u5bfc TFL1 [23]\uff0c\u8bba\u6587\u5c06 SOC1\u3001AGL24 \u548c XAL2 \u7684\u6d3b\u52a8\u805a\u5408\u5230 SAX \u4ee3\u7406\u4e2d\uff0c\u4e00\u8d77\u5145\u5f53 TFL1 \u6b63\u5411\u8c03\u8282\u5668[9]\u3002\u7531\u6b64\u521b\u5efa\u4e86 SALT\u57fa\u56e0\u8c03\u63a7\u7f51\u7edc<\/strong><\/span>\uff08\u56fe7\uff09<\/span>\uff0c\u7531\u8fd9\u56db\u4e2a\u8c03\u8282\u5668\u3001\u751f\u957f\u7d20Auxin<\/strong> [24]\u548c\u4ee3\u8868\u6210\u82b1\u6fc0\u7d20\u7684F\u503c<\/strong>\u7ec4\u6210\u3002\u540e\u8005\u662f\u4e00\u79cd\u8bf1\u5bfc\u82b1\u7684\u4fe1\u53f7\uff0c\u5f53\u690d\u7269\u8001\u5316\u6216\u66b4\u9732\u4e8e\u8bf1\u5bfc\u5f00\u82b1\u7684\u73af\u5883\u6761\u4ef6\u65f6\u8be5\u4fe1\u53f7\u4f1a\u589e\u52a0 [25,26]\u3002\u6700\u540e\u7814\u7a76\u8fd8\u5728\u7f51\u7edc\u4e2d\u6dfb\u52a0\u4e86\u4e00\u4e2aTFL1 \u7684\u65e9\u671f\u8fc7\u6e21\u6291\u5236\u56e0\u5b50\u4ee3\u7406<\/span>\uff08eREP\uff09<\/span>\u3002<\/span><\/section>\n
     <\/span><\/section>\n
     <\/span><\/section>\n
    <\/section>\n
    \u56fe7\uff1aSALT GRN \u7f51\u7edc\u548c\u56db\u4e2a\u8c03\u8282\u5668<\/span><\/section>\n
     <\/span><\/section>\n
    SALT\u7f51\u7edc\u7684\u771f\u5b9e\u57fa\u56e0\u8868\u8fbe\u6982\u62ec\u4e86\u5728\u5206\u5b50\u5c42\u6b21\u4f1a\u9a71\u52a8\u690d\u7269\u5206\u751f\u7ec4\u7ec7\u6210\u4e3a\u4f55\u79cd\u547d\u8fd0\u3002\u7136\u800c\uff0c\u6700\u7ec8\u690d\u7269\u7ed3\u6784\u4e0d\u4ec5\u53d6\u51b3\u4e8e\u5206\u751f\u7ec4\u7ec7\u7684\u7c7b\u578b\uff0c\u8fd8\u53d6\u51b3\u4e8e\u5f62\u6001\u52a8\u529b\u5b66\u53c2\u6570<\/strong>\uff0c\u5305\u62ec\u51b3\u5b9a\u901a\u8def\u7684\u5206\u5b50\u9608\u503c\u3001\u5668\u5b98\u751f\u957f\u901f\u7387\u3001\u5206\u751f\u7ec4\u7ec7\u5f00\u59cb\u751f\u6210\u5668\u5b98\u7684\u5ef6\u8fdf\u4ee5\u53ca\u5668\u5b98\u751f\u4ea7\u901f\u7387<\/strong>\uff0c\u8fd9\u4e9b\u90fd\u662f\u72ec\u7acb\u8c03\u8282\u7684\u3002LFY \u548c ap1 cal \u7a81\u53d8\u4f53\u5b9e\u4f8b\u8bf4\u660e\u4e86\u8fd9\u4e00\u70b9\uff0c\u5b83\u4eec\u5177\u6709\u76f8\u540c\u7684GRN\u8f93\u51fa<\/strong><\/span>\uff08\u56fe8C\uff09<\/span>\uff0c\u4f46\u5177\u6709\u660e\u663e\u4e0d\u540c\u7684\u7ed3\u6784\u548c\u5f62\u6001<\/strong>[8,29]\u3002<\/span><\/section>\n
     <\/span><\/section>\n
    \u56e0\u6b64\uff0c\u6700\u7ec8\u690d\u7269\u82b1\u5e8f\u7ed3\u6784\u6e90\u4e8e\u82b1\u57fa\u56e0\u8c03\u63a7\u7f51\u7edc (GRN) \u548c\u5f62\u6001\u52a8\u529b\u5b66\u53c2\u6570\u4e4b\u95f4\u590d\u6742\u7684\u76f8\u4e92\u4f5c\u7528<\/strong>\u3002\u4e3a\u7814\u7a76\u8fd9\u79cd\u76f8\u4e92\u4f5c\u7528\u5728\u62df\u5357\u82a5\u4e2d\u662f\u5982\u4f55\u8fd0\u4f5c\u7684\uff0c\u8fd8\u9700\u8981\u7ed3\u5408\u52a8\u529b\u5b66\u8fdb\u884c 3D\u690d\u7269\u8ba1\u7b97\u5efa\u6a21\u3002<\/span><\/section>\n
     <\/span><\/section>\n
    <\/section>\n
    \u56fe8\uff1aSALT\u7f51\u7edc<\/span><\/section>\n
     <\/span><\/section>\n

    \uff08\u56fe8 SALT GRN\u6a21\u578b\u53ca\u5b9e\u9a8c\u9a8c\u8bc1 (A)SALT GRN\u7f51\u7edc\u7ed3\u6784 (B)SAX\u3001AP1\/CAL\u3001LFY\u548cTFL1\u5728 WT \u548c ap1 cal \u7a81\u53d8\u4f53\u7684\u5206\u751f\u7ec4\u7ec7\uff08SAM\uff09\u548c\u4fa7\u539f\u57fa\u4e2d\u7684\u5df2\u77e5\u8868\u8fbe\u6a21\u5f0f\u3002\u95ee\u53f7\u8868\u793a\u6a21\u578b\u7684\u9884\u6d4b\u8868\u8fbe\u6a21\u5f0f\u3002(C)\u5728SAM\uff08\u4f4e\u751f\u957f\u7d20\uff09\u548c\u4fa7\u5206\u751f\u7ec4\u7ec7\uff08\u9ad8\u751f\u957f\u7d20\uff09\u4e2d\u4e0d\u540cF\u503c\u4e0b\u6a21\u578b\u7684 WT\u3001tfl1\u3001ap1 cal \u548c lfy \u7a33\u6001\u3002\u9884\u6d4b\u7684 WT \u548c\u6240\u6709\u7a81\u53d8\u4f53\u5206\u751f\u7ec4\u7ec7\u7684\u9057\u4f20\u7279\u6027\u4e0e\u5b9e\u9a8c\u89c2\u5bdf\u5230\u7684\u8868\u578b\u4e00\u81f4 (D)\u9ad8F\u503c\u4e0b\u4fa7\u539f\u57fa\u57fa\u56e0\u8868\u8fbe\u7684\u65f6\u95f4\u6a21\u62df \uff08E&F\uff09SOC1:GFP\uff08\u767d\u8272\/\u6d45\u84dd\u8272\u4fe1\u53f7\uff09\u57fa\u56e0\u62a5\u544a\u5728WT(E)\u548c ap1-7 cal-1 \u7a81\u53d8\u4f53(F)\u4e2d\u7684\u82b1\u5e8f\u8868\u8fbe\uff09<\/span><\/p>\n


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    3. \u5f62\u6001\u52a8\u529b\u5b66\u548cGRN\u7ed3\u5408\u7684<\/strong><\/strong><\/p>\n

    \u591a\u5c3a\u5ea6\u751f\u6210\u6a21\u578b<\/strong><\/strong><\/p>\n<\/section>\n

    \n

    <\/section>\n

    <\/section>\n<\/section>\n<\/section>\n<\/section>\n<\/section>\n<\/h3>\n

    <\/span><\/section>\n
    \u57fa\u4e8eSALT GRN\uff0c\u4f5c\u8005\u5c06L\u7cfb\u7edf<\/strong><\/span>\uff08L-System\uff09<\/span>\u96c6\u6210\u4e00\u4e2a\u52303D\u8ba1\u7b97\u56fe\u5f62\u6a21\u578b\uff1a\u7531\u5851\u9020\u5730\u9762\u690d\u7269\u7ed3\u6784\u7684\u56db\u79cd\u5668\u5b98\u6784\u6210\uff1a\u5206\u751f\u7ec4\u7ec7<\/strong><\/span>\uff08meristems\uff09<\/span>\u3001\u8282\u95f4<\/strong><\/span>\uff08internodes\uff09<\/span>\u3001\u82b1\u6735<\/strong>\u548c\u53f6\u5b50<\/strong><\/span>\uff08\u56fe9\uff09<\/span>\u3002\u6bcf\u4e2a\u5206\u751f\u7ec4\u7ec7\u7684\u72b6\u6001\u6807\u8bc6\uff08\u690d\u682a\u3001\u82b1\u5e8f\u548c\u82b1\u6735\uff09<\/strong>\u7531 GRN \u7a33\u6001 State(t) \u51b3\u5b9a\uff0c\u6bcf\u4e2a\u65f6\u95f4\u6b65\u7684\u8ba1\u7b97\u4e3a\u5206\u751f\u7ec4\u7ec7\u5148\u524d\u72b6\u6001\u548c\u5916\u90e8\u56e0\u7d20\uff08\u690d\u7269\u751f\u957f\u7d20auxin \u548c F\uff09\u7684\u51fd\u6570<\/strong>\u3002GRN\u6a21\u578b\u5b9e\u73b0\u4e3a\u5355\u5ba4\u5e38\u5fae\u5206\u65b9\u7a0b<\/strong> <\/span>\uff08\u89c1\u8bba\u6587\u8865\u5145\u6750\u6599\uff09<\/span>\u3002<\/span><\/section>\n
     <\/span><\/section>\n
    <\/section>\n

    \u56fe9\uff1a\u62df\u5357\u82a5\u591a\u5c3a\u5ea6\u53d1\u80b2\u6a21\u578b\u7684\u793a\u610f\u56fe\u3002\u6bcf\u4e2a\u5206\u751f\u7ec4\u7ec7\u72b6\u6001\u7531\u4fe1\u53f7\u6c34\u5e73\uff08\u751f\u957f\u7d20 auxin\uff0cF\uff09\u548cGRN\u7a33\u6001\u6784\u6210\u3002\u5728\u65f6\u95f4t\uff0c\u690d\u7269\u7531\u4e00\u7cfb\u5217\u5668\u5b98\u7ec4\u6210\u7684\uff08\u5de6\uff09\uff0c\u5728\u65f6\u95f4t+\u0394t\uff08\u53f3\uff09\uff0c\u8be5\u6a21\u578b\u66f4\u65b0\u6bcf\u4e2a\u5206\u751f\u7ec4\u7ec7\u7684\u4fe1\u53f7\u6c34\u5e73\u548cGRN\u72b6\u6001\u3002\u7a33\u6001\u5b9a\u4e49\u4e86\u7528\u4e8e\u8ba1\u7b97\u4fa7\u539f\u57fa\u5206\u751f\u7ec4\u7ec7\u7684\u6807\u8bc6\uff08\u690d\u682a\u3001\u82b1\u5e8f\u6216\u82b1\u6735\uff09\u3002\u7eff\u8272\u6570\u5b57\u8868\u793a\u5206\u751f\u7ec4\u7ec7\u7684\u987a\u5e8f<\/span><\/p>\n

     <\/span><\/section>\n
    \u8be5\u6a21\u578b\u8fd8\u5305\u542b\u63cf\u8ff0\u5668\u5b98\u751f\u957f\u52a8\u6001\u7684\u89c4\u5219<\/strong><\/span>\uff08\u8282\u95f4\u548c\u53f6\u7247\u4f38\u957f\u3001\u82b1\u7684\u751f\u957f\u3001\u5668\u5b98\u4ea7\u751f\u901f\u5ea6\u548c\u751f\u957f\u521d\u59cb\u5ef6\u8fdf\uff09<\/span>\u3002\u6a21\u62df\u690d\u7269\u4ece\u4e00\u4e2a\u5355\u4e00\u690d\u682a\u5206\u751f\u7ec4\u7ec7\u5f00\u59cb\uff0c\u6839\u636eGRN\u3001\u5f62\u6001\u52a8\u529b\u5b66\u89c4\u5219\u548cF\u7684\u8f93\u5165\u503c\uff0c\u4e0d\u65ad\u91cd\u590d\u4ea7\u751f\u65b0\u7684\u5668\u5b98\u3002\u901a\u8fc7\u5728\u5408\u7406\u8303\u56f4\u5185\u7684\u503c\u8c03\u6574GRN\u548c\u5f62\u6001\u52a8\u529b\u5b66\u53c2\u6570\uff0c\u4f5c\u8005\u6210\u529f\u5730\u6821\u51c6\u4e86\u6a21\u578b\uff0c\u4ea7\u751f\u4e86 WT \u548c lfy \u690d\u682a\u53ca tfl1 \u7a81\u53d8\u4f53<\/strong><\/span>\uff08\u56fe13B-D\uff0cmovie S1&S2\uff09<\/span>\u7684\u771f\u5b9e\u7ed3\u6784\u548c SAX \u4e0d\u5f00\u82b1\u7a81\u53d8\u4f53\u8868\u578b\u3002<\/span><\/section>\n

    <\/span><\/section>\n